Background
Group A Streptococcus (GAS) causes millions of infections annually. Interventions can be complicated by the high strain diversity often observed in disease-endemic settings. It is not well understood how high strain diversity is maintained in these populations given that strains compete with each other both directly (within an individual host) and indirectly (via host immunity). Several previous studies have investigated how indirect competition affects the prevalence and diversity of strains. However, these studies tend to make simplifying assumptions about the role of direct competition, such as virulence and metabolic type, that may occur within hosts. There is little data available to determine the validity of these assumptions and there is a need to clarify how sensitive model outputs are to these assumptions.
Methods
In this study, we develop an agent-based model which enables us to compare the epidemiological dynamics of a multi-strain pathogen under different assumptions about direct competition between strains.
Results
Our results suggest that while direct and indirect competition can each decrease strain diversity when they act in isolation, they may increase strain diversity when they act together. This indicates that GAS modelling studies should examine sensitivity to assumptions about direct competition. Ongoing model calibration using longitudinal endemic GAS surveillance data is being used to assess the effect of direct competition on population-level GAS strain diversity.
Conclusions
Omitting considerations of direct competition can lead to inaccurate estimates of the likely effectiveness of control strategies as changes in strain diversity shift the level of direct strain competition.